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Measurement of Lipid Peroxidation Lipid peroxidation level in the brain cell homogenate was quantified using the thiobarbituric acid reactive substances (TBARS) as previously described (Buege and Aust, Measurement of Protein Carbonyl Content Protein carbonyls were used as a measure of protein oxidation, in brain cell homogenates using DNPH as previously described (Levine et al., 2000) and using the BMG Fluostar Optima multimode plate reader (NY, USA)